“…During base flow, headwater streams act as surficial expressions of groundwater conditions, providing observable spatiotemporal information of groundwater storage within catchments (e.g., Bencala et al, 2011;Biswal & Nagesh Kumar, 2013;Godsey & Kirchner, 2014;Kirchner, 2009;Shaw et al, 2017;Whiting & Godsey, 2016). In rain-dominated climates, all stream flow during these dry periods must come from water stored belowground, typically in the form of slowly draining groundwater that is locally sourced from adjacent hillslopes or is derived from regional groundwater systems that may cross local hillslopes or topographic watershed divides (e.g., Broda et al, 2012Broda et al, , 2014Clark et al, 2009;Frisbee et al, 2016;Gleeson & Manning, 2008;McNamara et al, 2011;Payn et al, 2012;Sheets et al, 2015;Tague & Grant, 2004;T oth, 1963;Troch et al, 2003;Welch & Allen, 2012). The hydraulic conductivity, geometry, and volume of this storage source should impact both the persistence and distribution of wetted channels in the dry season.…”