“…Although translational direction signals are encoded in V1 (Hubel and Wiesel, 1968), MT (also known as V5; Zeki, 1974;Maunsell and Van Essen, 1983;Albright et al, 1984), and medial superior temporal area (MST; Zeki, 1980;Tanaka et al, 1986), neural correlates of global complexflow motion are first encountered in the dorsal portion of MST (MSTd; Saito et al, 1986;Graziano et al, 1994;Lagae et al, 1994;Smith et al, 2006;Britten, 2008). MSTd neurons have been hypothesized to integrate local translational motion signals from early visual cortices into global complex-flow motion perceptions (Wurtz and Duffy, 1992;Warren and Saunders, 1994;Royden, 2002;Layton et al, 2012;Mineault et al, 2012;Layton and Fajen, 2016;Yu et al, 2018); however, the details remain elusive.…”