“…CNNs can complement existing methods for morphological studies such as morphological matrix ( Dai, Korn & Song, 2021 ), landmark ( Bazzi et al, 2018 ), fractal dimensions ( Wiese et al, 2022 ), ornamentation index ( Miao et al, 2022 ), conch properties ( De Baets, 2021 ), and 3D morphological methods ( Klinkenbußet al, 2020 ) and provide new perspectives for studying the morphological evolution of fossils in the future. Geometric morphometry requires the extraction of fossil features by labelling manually and performing descending operations ( e.g.…”