A new theory, called the tilt constancy theory, claims that the Ponzo illusion is caused by the misperception of orientation induced by local visual cues. The theory relates the Ponzo illusion-along with the Zöllner, Poggendorff, Wündt-Hering, and cafe wall illusions-to the mechanisms that enable us to perceive stable orientations despite changes in retinal orientation or body orientation. In Experiment 1, the magnitude of the misperception of orientation was compared with the magnitude of the Ponzo illusion. In Experiment 2, predictions of the tilt constancy theory were compared with accounts based on (1) low spatial frequencies in the image, (2) memory comparisons (pool-and-store model), and (3) relative size judgments. In Experiment 3, predictions of the tilt constancy theory were tested against predictions of the assimilation theory of Pressey and his colleagues. In the final experiment, the orientation account was compared with theories based on linear perspective and inappropriate size constancy. The results support the tilt constancy theory.