“…Behaviors can be measured as instantaneous events by frequency (number of occurrences) or as states (time spent performing behavior, Altmann, 1974), or contextualizing events within states, when several events are clustered within a state (Tardin, Pinto, Alves, & Simão, 2014). Because there are no known ethograms, activity budgets, or behavioral assays for this species, and because we had high definition videos that could be repeatedly rescored, we elected to record states by four measures: the number of times each behavior was recorded (discrete events), total duration of all discrete events, the mean duration of each behavior, and the percentage of the behavior across the focal period.…”