The paper investigates the seismic response of square tunnels in sand by means of dynamic centrifuge testing and numerical analysis. A series of dynamic centrifuge tests conducted at the University of Cambridge on a square aluminium model tunnel embedded in dry sand, are initially presented. The tests, which were designed in order to investigate the seismic response of flexible tunnels, are analyzed numerically be means of full dynamic analysis of the coupled soil-tunnel system, using different soil and soil-tunnel interface models. Numerical predictions are compared to the experimental data, in order to better understand the response mechanism and validate the numerical modelling. The validated numerical models are then used to investigate the effect of the lining rigidity on the soil-tunnel system dynamic response. The experimental and numerical results reported herein, indicate a nonnegligible rocking deformation mode for the tunnels coupled with racking distortion during seismic shaking. The significant effects of the lining rigidity, soil-tunnel interface characteristics and soil yielding on the dynamic earth pressures and the shear stresses developed around the perimeter of the tunnel, as well as on the dynamic lining forces, are also reported and discussed. Response to Reviewers Comments The Authors gratefully acknowledge the constructive criticism of the Reviewers and their useful comments and suggestions, which helped to further improve the clarity and quality of the paper. Major modifications in the revised manuscript are highlighted in yellow. Below are the Authors' replies to individual comments: Reviewer #1: Reviewer's comment: The paper studies the seismic response of square tunnels combing experimental and numerical methods. The numerical analyses are validated against centrifuge-model tests, and the validated models are used to conduct a parametric study. The paper is well written and the subject is of interest to the readership of the journal. The paper should be published after the following drawbacks are addressed: 1) The beginning of the introduction gives the impression that tunnels are very sensitive to seismic shaking. Although there are cases of severe damage or collapse of tunnels in strong earthquakes, as correctly mentioned by the authors, tunnels are in general not so sensitive to seismic shaking. The introduction should be revised, so as not to give a wrong impression to the readers. Authors' reply: The Authors agree with the Reviewer. Tunnels were found less vulnerable to seismic shaking compared to above ground structures during recent strong earthquakes. To avoid any misunderstanding the introduction has been revised as follows: Pages: 1-2, lines: 31-33, 1-2 "Underground structures and tunnels behaved better than above ground structures during recent strong earthquakes. However, several cases of severe damages to total collapse have been reported in the literature (e.g. [27, 53]). Shallow embedded structures in soft soil were found more vulnerable to seismic shaking, while the vuln...