“…The microhylid subfamily Cophylinae Cope, 1889, endemic to Madagascar, is today recognised as possessing eight genera according to Scherz et al (2016a): Plethodontohyla Boulenger, 1882, Rhombophryne Boettger, 1880, Stumpffia Boettger, 1881, Madecassophryne Guibé, 1974, Platypelis Boulenger, 1882, Anodonthyla Müller, 1892, Cophyla Boettger, 1880 and Anilany Scherz, Vences, Rakotoarison, Andreone, Köhler, Glaw & Crottini, 2016. Recent studies (Vieites et al 2009, Perl et al 2014 have provided DNA barcoding reference sequences for almost all described and undescribed frog species of Madagascar and have highlighted the presence of a large taxonomic gap in the Cophylinae, which currently comprises over 100 described species (Scherz et al 2016a, AmphibiaWeb 2018, but which has an estimated further > 40 species yet to be formally described.…”