“…Li et al (2012) showed that the freshwater eelgrass Vallisneria natans grown in silt and clay had greater height, more ramets and leaves, and greater biomass accumulation compared to V. natans grown in pebble and gravel substrates. Healthy eelgrass beds are found in substrates with wide range of silt and clay content (1-56%; Imao and Fushimi, 1985;Keser et al, 2003;Koch, 2001;Laugier et al, 1999;Tamaki et al, 2002). Although, a high silt-clay content can create a reducing environment due to decreased exchange of pore water with the water column (Huettel and Rusch, 2000;Koch, 2001).…”