“…In an isotropic environment, active colloids of both living and inanimate types move along random directions unless their trajectories are biased by gradients of chemicals, temperature, or other cues [5][6][7]. Liquid crystals, used as a medium for active colloids, offer a much higher control level over the microscale dynamics thanks to their longrange orientational order [4,8,9]. In particular, by designing patterns of the nematic director n (n ≡ − n, n2 1) that specifies the preferred direction of molecular orientation [10], one can command the polarity and geometry of propulsion trajectories [11][12][13][14][15][16][17][18], mediate transitions from individual to collective modes of propulsion [17] and control the spatial distribution of microswimmers [14,15,17].…”