“…Chromosome studies are necessary to improve our knowledges on intra-and inter-specific as well as intraand inter-specific relationships in the most evolutionary advanced Asteraceae (e.g., Volkova and Boyko 1986; Vallès et al 2005). Studies of chromosome counts in several species of the genus were reported (e.g., ValdesBermejo et al, 1976, Brullo et al, 1977Bir et al, 1979;Bir et al, 1980;Chikkannaiah et al, 1982;Snogerup et al, 1985;Luque et al, 1991;Mejías et al, 1993;Razaq et al, 1994;Malallah et al, 1999;Kamel. 2004).…”