“…Pinus mugo has recently expanded into surrounding grassland or heath in many areas of the Carpathians (Solár & Janiga, 2013; Švajda, Solár, Janiga, & Buliak, 2011), Apennines (Calabrese et al, 2018; Dai et al, 2017), Alps (Dullinger et al, 2003) and Sudetes (Treml, Wild, Chuman, & Potůčková, 2010). Newly established individuals serve as islands that might quickly spread further by growth and clonal reproduction (Špinlerova, 2015; Wild & Winkler, 2008) and gradually overgrow alpine grasslands (Treml et al, 2010), which are rich in biodiversity (Kašák, Mazalová, Šipoš, & Kuras, 2015). In this study, we aimed to test the significance of individual establishment stages for subalpine shrubs, as represented by Pinus mugo .…”