Abstract. This paper, prepared for the forthcoming The Social Economics Handbook (Jess Benhabib, Alberto Bisin and Matthew Jackson, editors, Elsevier Press), surveys recent work on the evolutionary origins of preferences. We are especially interested in the circumstances under which evolution would push preferences away from the self-interested perfectly-rational expected utility maximization of classical economic theory in order to incorporate environmental or social considerations. * We thank Andrew Postlewaite for helpful discussions and comments.