U-Pb SHRIMP ages obtained in zircons from the Sotosalbos and Toledo anatectic complexes in Central Spain give new constraints to the evolution of the inner part of the Hercynian Iberian belt. Pre Hercynian ages in zircons from the Sotosalbos com plex (-464 Ma) are well preserved and reveal that an age diversity of the Lower Paleozoic magmatism in the area exists, as previous data on westernmost or thogneisses yield significant older ages. Zircon ages in the pelite-derived granites from the Toledo complex also show an important N eoproterozoic age compo nent which points to a metasedimentary protolith deposited maximally 560 Ma ago. Younger zircon populations in both complexes at -330 Ma in the Sotosalbos region and -317 Ma in the Toledo com plex indicate an important diachronism between the anatectic processes in both areas but also that these processes are mainly unrelated to the generation of the later Hercynian granite batholith of Central Spain, which could be of deeper crustal derivation. In addition, as migmatization occurred late in the metamorphic cycle, after peak conditions were at tained, the age of anatexis is younger than the age of the main Hercynian metamorphic event, which still is not well constrained.