“…Ectomycorrhizal fungi synthesize oxalic acid (Lapeyrie, Chilvers & Bhem, 1987), which is able to chelate Al to a non-toxic form in growth media like other organic acids (Jones, 1961;Sivasubramaniam & Talibudeen, 1971;Rajan et al, 1981). Many sugars such as glucose, fructose, galactose, maltose, arabinose and rhamnose, produced and secreted by roots and microbes into their growth media, form metallo-organic complexes with Al (Bradley & Sieling, 1953).…”