“…These insects have been used in ecological studies because of their ease of culture, as well as in research on parasite-induced host behaviors because of their role as intermediate hosts for helminths. Moreover, they have been studied in efforts to develop control methods (Shostak and Smyth, 1998;Yan et al, 1998;Pai and Yan, 2003;Arnaud et al, 2005). Flour beetles also represent interesting host-parasite systems in that all or most of the species, including both adults and larvae, are infected with gregarine parasites (Apicomplexa: Conoidasida: Gregarinasina: Eugregarinorida), and the presumed relationship among hosts provides a tractable laboratory model to explore experimentally the evolutionary constraints on host specificity.…”