“…However, understory plant species have heterogeneous abundance and distribution patterns, which depends on the overstory species and structure (e.g., site quality of the stands or gap presence) (e.g., Warner and Harper, 1972;Veblen et al, 1977Veblen et al, , 1979Goldblum, 1997;Hutchinson et al, 1999;Damascos and Rapoport, 2002;Small and McCarthy, 2005;Fahey and Puettmann, 2008), as well as of micro-environmental and stand conditions (e.g., Huebner et al, 1995;Thomas et al, 1999;Palmer et al, 2000;Fraterrigo et al, 2009). Natural and anthropogenic disturbances in forest ecosystems affect these conditioning factors and modify understory diversity and distribution (Chen et al, 1993;Ohlson et al, 1997;Jenkins and Parker, 1999;Økland, 2000;Martínez Pastur et al, 2010). Therefore, herbaceous vegetation could act as a useful indicator of forest sustainability, ecosystem health and conservation status Lindenmayer et al, 2000;Dale et al, 2002).…”