“…For example, anatomical characters, especially of leaves, have been extensively used in the systematics of several plant families, including Bromeliaceae (De Faria et al, 2012), Myrtaceae (Al-Edany and Al-Saadi, 2012), Amaryllidaceae (Lin and Tan, 2015), Malpighiaceae (Araújo et al, 2010) and Asteraceae (Castro et al, 1997;Milan et al, 2006;Adedeji and Jewoola, 2008;Bombo et al, 2012;Akinnubi et al, 2014;Rojas-Leal et al, 2017;Lusa et al, 2018). Particularly in Asteraceae, leaf architecture and anatomical characters are extremely diverse (Bombo et al, 2012;Rojas-Leal et al, 2014, 2018 and have been studied from an ecological (Bercu et al, 2012;Moroney et al, 2013;Rivera et al, 2017;Ferraro and Scremin-Dias, 2018), physiological (Bondarev et al, 2003;McKown and Dengler, 2007;Santiago and Kim, 2009), and medical perspective (Cambi et al, 2006;Hulley et al, 2010;García-Sánchez et al, 2012).…”