Comparing life histories of shrews and rodents. [In: Theriology at the turn of a new century. J. Gliwicz, ed]. Acta Theriologica 47, Suppl. 1: 185-208.Small insectivores and rodents, despite similarities in body size and attributes scaling to body size, exhibit significant differences in other properties, including many life history traits. In this article major differences between life history traits of the two taxa are reviewed, with an indication of contrasting selection pressures related to somewhat different body size, as well as to differences in metabolic rates, diet and exposure to predation. Additionally, since the life history differences between small mammals are particularly well pronounced in highly seasonal habitats, the winter ecology of shrews and rodents is compared. Finally, the two different reproductive strategies typical for soricine shrews and small nonhibernating rodents, are presented. In conclusion, it is proposed that the reproduction delayed to the second calendar year of life in shrews is the result of selection for traits ensuring successful survival in winter, a period that is more perilous for shrews than for rodents. In rodents, in contrast, opportunistic reproduction is the most prominent characteristic which also helps to maximize their reproductive output. This ability for high reproduction seems to be the main antipredatory measure selected for in rodent evolution.