This paper examines the ongoing significance of locative media and mobilegenerated geocoded data, including their increasing integration into the core functionalities and business objectives of large social media and search services. In this article, I take a 'communicative ecologies' approach to explore how location-based services function as a dynamic system, with a fluid and shifting structure or set of relations. The evolution of the mobile social networking and search and recommendation service Foursquare provides a striking example of such a system. In the first half of the paper, I explore the company's still-evolving business model, and their intricate corporate relationships with other key search, recommendation, and social media firms. In the second half, I trace how this dynamic engagement is also at play in the end uses of Foursquare and other location applications.