RESEARCH M aize provides about 30% of the food calories for approximately 4.5 billion people in 94 developing countries (Shiferaw et al., 2011); globally, however, 63% of maize is used for livestock feed. Maize kernel provides 365 kcal 100 g-1 ; it is composed of approximately 73% starch, 10% protein, and 5% oil, with the remainder made up of fiber, vitamins, and macro-and micronutrients (Wang et al., 2012). The composition is largely controlled by the genetics of the endosperm sink, the maternal plant parent, and the environment (Nuss and Tanumihardjo, 2010). Since the 1920s successively released varieties having greater yield have had more starch and less protein contents (Scott and Blanco, 2009). However, kernel composition has been successfully manipulated by selection and may affect the nutritional value of the grain (Welch and Graham, 2004) and the value of end-use products (Ortiz-Monasterio et al., 2007). Farmers and the food and feed industries will continue to demand high-yielding maize varieties with enhanced nutritional and industrial qualities. Therefore, one of the most important