“…Although the difference between WUEe and WUEc can be effectively understood at the observation level (Wang, Li, & Tan, 2020), it is difficult to spatiotemporally distinguish the water consumption of ecosystems and canopies when they are quantitatively stripped at a large scale (Bai et al, 2020). Therefore, studies on WUE have usually considered only WUEe (Yan et al, 2021). Early WUE studies mainly focused on crop leaves or individual levels (Donovan et al, 2007) and gradually extended to farmlands (Wang et al, 2018), grasslands (Wang, Ma, Li, & Yuan, 2020), forests (Jiang et al, 2019) and other ecosystems (Driscoll et al, 2020).…”