“…Consistent amongst the reviewed articles was that naturalistic exhibits (i.e. exhibits designed to be aesthetically appealing and mimic natural habitats [58]) were associated with more positive welfare perceptions [11, 16, 18, 19, 29, 30, 32, 35, 37, 39, 40, 42–44, 46, 47, 51, 52, 62, 64, 65, 70, 88, 95–98, 105, 109, 119–129]. However, naturalistic enclosures may impinge on expression of natural behaviour as naturalistic aesthetics do not always equate to usable space [95, 130].…”