“…between quadrats and between sites) as the most relevant for the variability of the distribution of almost all size classes of Paracentrotus lividus, though the distribution of the largest individuals varied considerably also between shores located some kilometers apart. The small-scale patchy distribution of present populations of sea urchins, particularly of relatively small individuals, agrees with previous studies which have demonstrated the main variability of the distribution of sea urchins at spatial scales from centimeters to meters rather than at larger scales (Hereu et al, 2004;Ceccherelli et al, 2009Ceccherelli et al, , 2011Addis et al, 2012;Bertocci et al, 2012b). Several biological and abiotic factors and processes, separately or possibly through complex interactions, can be responsible for such patterns, including competition (Guidetti et al, 2004), predation (Sala and Zabala, 1996;Hereu et al, 2005;Guidetti, 2006), recruitment (Tomas et al, 2004;Rius and Zabala, 2008) and topographic complexity (Benedetti-Cecchi and Cinelli, 1995;Barnes and Crook, 2001b;Hereu et al, 2005).…”