Ptychoptera delmastroi sp. n. (Diptera: Ptychopteridae) from Italy

Helen’s twins in the Balkans: discovery of two new Paraptychoptera Tonnoir, 1919 species closely related to P. helena Peus, 1958, with systematic revision of the “lacustris” group (Diptera, Ptychopteridae)

“…The study of P. helena was based on a paratype male from the Zoological Research Museum Alexander Koenig (ZFMK), Bonn, Germany, Museum-Id ZFMK-DIP-00015966. Krzeminski & Zwick, 1993, P. delmastroi Zwick & Stary, 2003, and P. ressli Theischinger, 1978 were not available during the present study, but the detailed morphological description of wing and male terminalia, based on Krzeminski and Zwick (1992) and Zwick and Starý (2003), were used to evaluate discrete details on male terminalia. A comprehensive morphological dataset of the world Ptychopteridae published recently by Fasbender (2014) was also used as a source for morphological details of the male terminalia in species where fresh material were not available to us during the study.…”

“…Later, the group was synonymised by Alexander (1927) and included between the more widespread Ptychoptera and, consequently, was ignored in important revisions and contributions to the European Ptychopteridae (Freeman 1959;Peus 1958;Zitek-Zwyrtek 1971;Deliné-Draskovits 1983;Krzeminski 1986;Rozkosny 1997;Krzeminski and Zwick 1993). Paraptychoptera was recovered only recently as a subgenus of Ptychoptera (Zwick and Starý 2003;Fasbender 2014) and also referred to by Ujvárosi et al (2011). Strong morphological and molecular support to monophyly of Paraptychoptera were added in a reference contribution of the world Ptychopteridae species (Fasbender 2014).…”

“…silvicola, Zwyrtek &Rozkosny, 1967 are restricted to Central or Southern Europe. Another five species, however, have a more restricted distribution: P. (P.) agnes Krzeminski & Zwick, 1993 is an endemic species that is described from the Pilis mountains, in the Transdanubian Mountains, Hungary (Krzeminski and Zwick 1993), P. (P.) delmastroi Zwick & Stary, 2003 from Villafranca municipality in the City of Turin, and Cantarana municipality (both are in the region of Piemonte), northern Italy (Zwick and Starý 2003) and Tunisia (Paramonov, 2013) while P. (P.) helena Peus, 1958 is known only from a single site, Oiti village, Mount Oeta, Phthioitis region, Greece (Peus 1958).…”

“…Among Paraptychoptera, the lacustris group was first proposed by Tonnoir (1919), as species having less developed, but recognisable, auxiliary sexual organs in the male sternite III, and was also referred to by Stubbs (1993) and Zwick and Starý (2003), and included the following species: P. lacustris, P. longicauda, P. paludosa. In the present paper we describe two new species of the Ptychoptera ( Paraptychoptera) lacustris from the South Balkans, and provide a key of the revised lacustris group, within which the two new species belong.…”

Global diversity of dipteran families (Insecta Diptera) in freshwater (excluding Simulidae, Culicidae, Chironomidae, Tipulidae and Tabanidae)

“…Paraptychoptera Tonnoir is a monophyletic endemic European clade and was assigned subgeneric rank within Ptychoptera, in order not to turn Ptychoptera s. str. paraphyletic (Zwick & Starý, 2003).…”