Downsized, highly boosted, GDI engines are becoming the preferred gasoline engine technology to ensure that increasingly stringent fuel economy and emissions legislation are met. The Ultraboost project engine is a 2.0 L in-line 4-cylinder prototype engine, designed to have the same performance as a 5.0 L V8 naturally aspirated engine but with reduced fuel consumption. It is important to examine Particle Number (PN) emissions from such extremely highly boosted engines to ensure that they are capable of meeting current and future emissions legislation. The effect of such high boosting on PN emissions is reported in this paper for a variety of operating points and engine operating parameters. The effect of engine load, air fuel ratio, fuel injection pressure, fuel injection timing, ignition timing, inlet air temperature, EGR level, and exhaust back pressure have all been investigated. It is shown that PN emissions increase with increases in cooled, external EGR and engine load, and decrease with increases in fuel injection pressure and inlet air temperature. PN emissions are shown to fall with increased exhaust back pressure, a key parameter for highly boosted engines. The effects of these parameters on the particle size distributions from the engine have also been evaluated. Significant changes to the particle size spectrum emitted from the engine are seen depending on the engine operating point. Operating points with a bias towards very small particle sizes were noted.