“…The Japanese Archipelago extends along a 3,000 km line from the northeast to the southwest, covering a wide range of climatic zones, from subarctic to subtropical. The distribution ranges of most plants in the Japanese Archipelago have repeatedly shifted following climatic changes during the Quaternary (Ikeda, Carlsen, Fujii, Brochmann, & Setoguchi, ; Kubota, Kusumoto, Shiono, & Tanaka, ; Yoshida, Kudo, Shimada, Hashizume, & Ono, ). Because their range shifts in the archipelago interacted with the neighboring continent of Asia as well as associated islands (Chiang et al, ; Fujii, Ueda, Watano, & Shimizu, ; Ikeda, Higashi, Yakubov, Barkalov, & Setoguchi, ; Lee, Lee, & Choi, ; Nakamura et al, ; Sakaguchi et al, ), elucidating the historical interaction with the continent and surrounding islands is necessary to understand the origin of Japanese flora.…”