“…1, when applied to biophysical models of photosynthesis and transpiration, closely matches the functional forms of empirical stomatal models, which themselves have extraordinary empirical support (2,5,7,27,34). For this reason, the WUEH is widely used to guard against out-of-sample behavior in the novel climates created in ecosystem models (35)(36)(37)(38)(39). Finally, several notable efforts have been made to extend WUEH to situations including soil moisture stress (11,12,28,33,(40)(41)(42) and competition (43) and to develop a theory of the value of λ over the short time scales during which it should be approximately constant and how λ should change over weeks, months, or seasons (9,11,12,28,32).…”