It is well established that the large-scale oceanic fronts associated with the Kuroshio Extension and the Gulf Stream anchor the entrance regions of the midlatitude atmospheric storm tracks of the Northern Hemisphere (e.g., Nakamura et al., 2004Nakamura et al., , 2008. It is also widely accepted that the SST anomalies associated with the ocean mesoscale eddies that form along the large scale oceanic fronts have a major effect on the atmospheric boundary layer (e.g., Chelton & Xi, 2010;Small et al., 2008;Xie, 2004). The deeper tropospheric effects of SST mesoscale variability, however, are less understood and have been the subject of ongoing research. For instance, Woollings et al. (2010) found a subtle, but significant effect on the midlatitude storm tracks in simulations with a limited area version of the Hadley Center's third generation atmospheric model.