“…Yet, despite their ubiquity, limpet species can sometimes be difficult to tell apart in the field (Simison and Lindberg, 2003;Burdi, 2015), at archaeological sites (Rogers and Weisler, 2020a) and in museum collections (Kuo and Sanford, 2013) owing to their highly variable shell morphologies and colour patterns (Nakano and Spencer, 2007). Even within species, shell features can vary according to substrate, size (age), population, and geographic region, sometimes resulting in distinct shell morphologies (Williams, 2017) and shapes (Rogers and Weisler, 2020b) To further complicate matters, shell erosion and encrusting symbionts can also impede visual identification. As a result, taxonomists frequently rely on using internal anatomical features, such as radular structure, as distinguishing characters (Simison and Lindberg, 1999).…”