U-Pb geochronologic analyses have been conducted on 205 individual detrital zircon grains from Cambrian through Devonian sandstones of the RobertsMountains allochthon in central Nevada. These strata were tectonically emplaced onto the Cordilleran margin during the mid-Paleozoic Antler orogeny, but their original depositional settings and provenance have been controversial. Our data, combined with previous detrital zircon studies, define four different age signatures for the eugeoclinal strata: (1) 690-7 15 and 1065-1350 Ma grains in a minor group of sandstones in the Upper Cambrian(?) Harmony Formation, (2) 1745-Comparison of these data with the detrital zircon reference for the Cordilleran miogeocline and with ages of basement provinces in cratonal North America indicates that sandstones in the lower Vinini and parts of the Harmony Formations were derived from 1.0-1.3,~1.43, and 1.6-1.8 Ga provinces of southwestern North America. In contrast, most older and younger units contain few grains of the appropriate ages to have come from the southwestern part of North America, and instead have strong similarities with the Peace River arch region of western Canada.We propose that detritus in most of the Harmony Formation was shed from off-shelf basement rocks exposed along the Canadian continental margin, perhaps as a western continuation of the Peace River arch or as extensional fault blocks. In contrast, detritus