Transtensional basins embedded in the San Andreas fault system of Southern California (United States) and northwestern Mexico are fi lled with sediment derived from the Colorado River, which drains a large area of the western U.S. interior. The sediment is rapidly buried, heated, and mingled with intrusions in the deep basins to form a new generation of recycled crust along the active plate boundary. Using a range of values for total basin depth, relative volume of mantle-derived intrusions, and composition of early rift deposits, the volume of Colorado River-derived sediment in the basins is bracketed between 2.2 and 3.4 × 10 5 km 3 , similar to the volume of rock that likely was eroded from the Colorado River catchment over the past 5-6 m.y. The volumetric rate of crustal growth by sedimentation is ~80-130 km 3 /m.y./km, comparable to growth rates in subductionrelated island arcs and slow seafl oor spreading centers. Sedimentary and basinal processes thus play a major role in crustal evolution and recycling in this setting, and may be important at other rifted margins where a large river system is captured following tectonic collapse of a prerift orogenic highland.