“…republication, systemmatic reproduction, or collective redistirbution of any portion of this article by photocopy machine, reposting, or other means is permitted only with the approval of The oceanography society. send all correspondence to: info@tos.org or Th e oceanography society, Po box 1931, rockville, Md 20849-1931 volcanism driven by hotspots or mantle plumes within Earth's crust (see Staudigel and Clague, 2010 as well as the discovery of a host of bacteria and archaea with novel anaerobic and aerobic metabolisms that take advantage of unique suites of energy sources (both autotrophic and heterotrophic) derived from the combination of geochemistry, temperature, and pressure at these sites (Karl, 1995;Nakagawa and Takai, 2008 (Gold, 1992;Whitman et al, 1998 Lavelle and Mohn, 2010).…”