Literature concerning dinosaur footprints or trackways exhibiting abnormal gait or morphology reflecting pathology (ichnopathology) is rare. We report on a number of Jurassic and Cretaceous occurrences of theropod footprints from western North America with unusual morphologies interpreted herein as examples of inferred pathologies, or ichnopathologies. The majority of ichnopathologies are primarily manifested in the digit impressions and include examples of swelling, extreme curvature, dislocation or fracture, and amputation. A number of occurrences are single tracks on ex situ blocks with substantial deformation (inferred dislocation or fracture), or absence of a single digit impression. Two occurrences are from in situ natural mould trackways, one of which is a lengthy trackway of a presumed allosauroid with no noticeable deformation of the digits or feet but with strong inward rotation of the left footprint toward the midline and a pronounced, waddling limp. The other is a tyrannosaurid trackway consisting of three footprints (one right, two left) with the two left prints exhibiting repetitive ichnopathology of a partially missing Digit II impression.Keywords Dinosaur tracks, Amputation, Antalgic gait, Compensated gait, Jurassic, Cretaceous, Ichnopathology, Laterality, Irregular gait, Limping gait, Shuffling gait, Waddling gait
INTRODUCTIONOver the course of the past few decades, there has been a marked increase in the literature of pathology of dinosaurs (e.g., Tanke and Rothschild, 2002 and references therein). However, the literature on inferred pathology from dinosaur tracks and trackways (ichnopathology) has not seen a corresponding increase. Reports on ichnopathology in dinosaurs have been infrequent, with perhaps less than one paper published per decade since the beginning of modern studies of dinosaur traces in the early 19th century. It is possible that reports of ichnopathology have not kept pace with reports of pathologies on osteological specimens due to the various influences of track-bearing substrate, the behavior of the trackmaker, as well as a range of taphonomic factors that make such interpretations difficult. The sparse publication record of ichnopathology may also be a reflection of the cautious approach ichnologists employ when interpreting phenomena related to tracks.Even with a restrained approach toward footprint and trackway interpretation, some dinosaur tracks and trackways exhibit morphologies that have defied explanation by conventional behaviors of the trackmaker, or by the physical and dynamic qualities of the track surface, both pre-and post-burial.This article discusses previous reports of dinosaur ichnopathology with reinterpretations where such is warranted. Several recent and new reports of dinosaur ichnopathologies are described, from sites in the Jurassic of China and the Cretaceous of North America. A number of criteria are proposed that will aid in the interpretation of ichnopathologies