Plant breeding is man-directed evolution. Plant breeders manipulate the genetic resources of a species, i.e., its germplasm, to produce plants that are of increased value to humanity. The same analogy applies to animal improvement programs. All of our major food crops and all of our domestic animals and their respective breeds, strains, or cultivars were developed by this process. Although humans have successfully manipulated the genetic resources of plants and animals for several thousand years, the science of genetics and breeding was not developed until this century. Breeding work on most forage crops did not began until the 1930's and initial work was focused on developing strains that had good establishment capability, persistence, high forage yields, and had good insect and disease resistance. These are essential attributes of forages (Burton, 1986). This initial breeding work resulted in the development of grasses such as 'Coastal' bermudagrass (Cynodon dactylon L.), 'Lincoln' bromegrass (Bromus inermis Leyss.), and 'Kentucky 31' tall fescue (Festuca arundinacea Schreb.). Limited animal evaluation was involved in the development of these cultivars. Breeding for improved forage quality was not an important research objective of most forage programs until the last 25 yr. Kneebone (1960) published a review of grass breeding and did not discuss breeding for improved forage quality. However, some earlier reviews on grass breeding including the extensive review of Hanson and Carnahan (1956) included minor sections on breeding for improved forage quality. It was not until the pioneering research of Dr. Glenn Burton and his associates at Tifton, GA demonstrated the economic value of improved digestibility in bermudagrass that breeding for forage quality became a major research objective of some grass breeding programs (Burton, 1972a; Chapman et aI., 1972).