S U M M A R YRecent dense deployments of portable digital seismographs have provided excellent control on earthquakes beneath the central North Island of New Zealand. Here we use a subset of the best-recorded earthquakes in an inversion for the 3-D Vp and Vp/Vs structure. The data set includes 39 123 P observations and 18 331 S observations from 1239 earthquakes and nine explosions. The subducted plate is imaged as a high Vp, low Vp/Vs feature. Vp within the mantle of the subducted slab is almost always >8.5 km s −1 , which requires the ca. 120 Myr slab to be unusually cold. The low Vp/Vs within the subducted plate closely parallels the lower plane of the dipping seismic zone. It most likely indicates fluid resulting from dehydration of serpentine in the slab mantle, and the earthquakes themselves are likely to be promoted by dehydration embrittlement. We identify a region with Vp < 8.0 km s −1 which coincides with the upper plane of the dipping seismic zone and extends to ca. 65 km depth with the subducted Hikurangi Plateau, which is about 17 km thick prior to subduction. The mantle wedge is generally imaged as a low Vp, high Vp/Vs feature. However, there are significant changes evident in the wedge along the strike of the subduction zone. The region where Vp is lowest (7.4 km s −1 ) and Vp/Vs is highest (1.87) occurs at 65 km depth, immediately west of the Taupo caldera. This region is best interpreted as a significant volume of partial melt, produced by the reaction of fluid released by dehydration of the subducted plate with the convecting mantle wedge. The region with lowest Vp, while paralleling the underlying dipping seismic zone, is located about 30 km from the upper surface of the zone. Material with Vp > 8.0 km s −1 directly above the dipping seismic zone can be interpreted as sinking, entrained with the motion of the subducted slab and forming a viscous blanket that insulates the slab from the high-temperature mantle wedge. Material in the overlying low Vp region can be interpreted as rising within a return flow within the wedge. The volcanic front appears to be controlled by where this dipping low Vp region meets the base of the crust. The thickness of the backarc crust also shows significant variation along strike. In the central Taupo Volcanic Zone (TVZ) the crust is ca. 35 km thick, while southwest of Mt Ruapehu the crust thickens by ca. 10 km. There is no significant low Vp zone in the mantle wedge in this southwestern region, suggesting that this thicker crust has choked off mantle return flow. The seismic tomography results, when combined with constraints on mantle flow from previous shear-wave splitting results, provide a plausible model for both the distribution of volcanism in the central North Island, and the exceptional magmatic productivity of the central TVZ.