“…The epidermis is a multifunctional tissue, the size and state of cells of which largely determine the plant's water balance, the intensity of CO 2 assimilation, respiration, which ultimately affects the nature of the production process. The histological features of the cells of the primary integumentary tissue are determined genetically, but their formation and functioning depend on many biological and environmental factors, therefore, in the qualitative and quantitative study of the epidermis, it is necessary to pay special attention to the influence of each of these factors not only separately, but also in their synergistic effect (Vasylevska, 2014;Weaver & Clements, 2008;Wilson, 2002;Wright & Westoby, 2001;Zlobin, 2004;Crawley, 2000;Didukh & Plyuta, 2004;Didukh & Shelyag-Sosonko, 2001;Didukh & Shelyag-Sosonko, 2003;Golubets, 2017;Grozdynskyi, 2013;Grytsyk, Neiko & Melnyk, 2016;Gulac, Marchenko, Kapitanenko, Kuris & Oleksenko, 2022;Gulac, Oleksenko, Kaluha, Kravchenko & Yukhymenko, 2022;Hajibagheri, 2003;Jones, 2012;Kazakov, 2000;Khrystova & Pyurko, 2007;Khrystova & Pyurko, 2009).…”