“…Beyond the human drivers and conservation context of park upstream watersheds considered here, future analyses need to explicitly consider the ecological benefits and buffering potentials of natural systems. While some of these variables -like the percentage of natural land cover types -will be inversely correlated with many of the landscape stressors (e.g., percentage urban, percentage agriculture), others related to landscape pattern (Riitters et al, 1995(Riitters et al, , 2007(Riitters et al, , 2009a(Riitters et al, , 2009b and habitat connectivity (Hilty et al, 2006;Theobald, 2006;Goetz et al, 2009;Galpern et al, 2011) will provide key management insights at local to regional scales.…”