“…Water-surface and energy gradients over riffles are typically steeper than over pools at low flows and generally converge as stage increases and relative roughness decreases (Leopold et al, 1964;Keller, 1971;Richards, 1976;Wohl et al, 1993). Similarly, mean velocity and shear stress are greater over riffles than pools at lows flows, but the reverse may occur at high flows where channel conditions promote that phenomenon (Keller, 1971;Carling, 1991;Clifford and Richard, 1992;Wilkinson et al, 2004;MacWilliams et al, 2006;Harrison and Keller, 2007). Variations in depth, velocity and water surface gradients between riffles and pools at low flows lead to distinct sediment sorting patterns that may even persist at high flows and affect channel hydraulics (Keller, 1971;Lisle, 1979).…”