“…They rely on the fundamental hypothesis that a causal relationship exists between the modern distributions of plants and the associated climates (Jackson and Williams, 2004, and references therein). These techniques can be divided into two types: (1) those based on plant assemblages -modern analogue technique (MAT) (Overpeck, 1985;Guiot, 1990), weighted averaging (WA) (ter Braak and Looman, 1986;ter Braak and van Dame, 1989;Birks et al, 1990), weighted averaging-partial least-squares regressions (WA-PLS) (ter Braak and Juggins, 1993), artificial neural networks (ANNs) (Malmgren et al, 2001) or regression trees (Salonen et al, 2012) -and (2) those based on plant distributions -mutual climatic range (MCR) (Atkinson et al, 1987;Sinka and Atkinson, 1999;Elias, 1997), the coexistence approach (CA) (Mosbrugger and Utescher, 1997;Utescher et al, 2014) or probability density functions (PDFs) (Kühl et al, 2002;Truc et al, 2013). These methods are fully detailed in Birks et al (2010).…”