“…Thus, unlike alluvial rivers, the location of sandbars in the geomorphically constrained Colorado River are fixed within 13 geomorphic reaches (Table 2). More than 550 debris fan complexes generate a distinctive suite of fluvial microhabitats, each with discrete grain sizes, soils, stage elevation relationships, inundation frequencies, and stage-zoned riparian vegetation (Turner and Karpiscak 1980;Schmidt and Graf 1990;Melis et al 1997). Analyses of flow regulation influences at a large suite of debris fan complexes revealed that flow regulation allowed extensive development of fluvial marshes, enhancing riverine plant species richness (Stevens et al 1995;Waring et al in press).…”