Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)

Abstract

This paper presents a reclassification of the north temperate taxa associated with the genus Staphylinus sensu lato, and comments on relevant subtribes of the tribe Staphylinini. The treatment contains the following: (1) a historical introduction to the past and recent classification of these genera; (2) a discussion of the characters used herein, many of them being unconventional and used for the first time, and many of which are illustrated either by SEM photomicrographs or by line drawings; (3) a key to the north temperate subtribes of the tribe Staphylinini; (4) a discussion of the diagnostic characters of each of the five relevant subtribes (Philonthina, Quediina, Anisolinina, Staphylinina, and Xanthopygina); (5) a key to the north temperate generic-and subgeneric-level taxa associated with the genus Staphylinus sensu lato; (6) type species information and a discussion of diagnostic characters for each taxon at the generic and subgeneric level; (7) a discussion of reasons for taxonomic and nomenclatural steps taken at both generic and specific levels; (8) a checklist of north temperate taxa; and (9) a list of the references mentioned in the discussions in the text (not including those associated with citations of scientific names).At the subtribal level, Craspedomerina, originally erected by Bernhauer (1911: 88) as Craspedomeri, is placed in synonymy with Philonthina. Triacrina, originally erected as Triacri by Bernhauer (1931: 84), is placed in synonymy with Xanthopygina (both syn. nov.).At the generic level, new concepts are presented for Staphylinus, Dinothenarus, Parabemus, Ocypus, Matidus, Pseudocypus, Tasgius, and Rayacheila.

Systematics and phylogeny of the Andean genera Leptopeltus Bernhauer and Leptopeltoides gen. nov. (Coleoptera: Staphylinidae), with biogeographical notes

“…HW -head capsule maximum width (measured at widest point) HL -length of head capsule, from anterior margin of frontoclypeus to neck constriction (along midline) PW -pronotum maximum width PL -pronotum length (along midline) EL -eye length (seen from above) TL -temple length (from the posterior margin of the eye to the nuchal groove; seen from above) NW -neck width S1 -first segment of hind tarsus length S5 -last segment of hind tarsus length EtL -elytra length at sides (from humerus to apex; seen from above) Etl -elytra length along suture Terminology mainly follows Blackwelder (1936) and Schillhammer (2000) for the external morphology, Smetana and Davies (2000) for the head capsule ridges and Smetana (1995) for characters associated to male and female genitalia.…”

Generic Revision of the Procirrina (Coleoptera: Staphylinidae: Paederinae: Pinophilini)

“…15, 16). Generally the Paederinae, including the Procirrina, lack the abundance of cephalic ridges described by Smetana and Davies (2000), but many paederines do have a transverse carina or ridge on the dorsal surface that extends onto the lateral surface. Smetana and Davies (2000: 5, 8) discussed and illustrated a similar carina that they referred to as the nuchal ridge.…”

“…Blackwelder (1936: figs. 22, 23) presents them as a pair of black spots and Smetana and Davies (2000: fig. 2) label them as fossae and they are shown, but unlabeled, in their figures 42 and 54.…”

Xanthopygoides niger Cameron, 1951 (Xanthopygina) belongs to the genus Philonthus Stephens, 1829 (Philonthina): systematic and nomenclatural changes for the African Staphylinini (Coleoptera, Staphylinidae, Staphylininae, Staphylinini)

“…Such features of Philonthus neoniger as smooth pronotal disk with only one puncture in each of two dorsal "rows", dense and rugose punctation of elytra and abdomen, and, especially, a rather peculiar shape of the aedeagus (Figs 3,4) allow it to be unambiguously placed in the subgenus Raucalius Tottenham, 1949. Members of the subtribe Xanthopygina, instead, are usually characterized by a complete postmandibular ridge; the inferior marginal line of pronotal hypomeron well separated from the superior marginal line, both lines continuing uninterrupted and separated along the prosternal margin. Also, unlike Philonthina, all members of Xanthopygina have a pair of empodial setae (Smetana and Davies 2000). (Cameron, 1948), comb.…”