“…Evidence for this has been shown both at the colony level (Arechavaleta-Velasco & Guzmán-Novoa, 2001;Guzmán-Novoa, Emsen, Unger, Espinosa-Montaño, & Petukhova, 2012;Mondragó n, Spivak, & Vandame, 2005;Rinderer et al, 2001) and through assays inoculating individual bees with mites (Aumeier, 2001;Büchler, Drescher, & Tornier, 1992;Fries, Huazhen, Wei, & Jin, 1996;Guzmán-Novoa et al, 2012;Peng, Fang, Xu, & Ge, 1987). These are reviewed in detail by Rinderer et al (2013), and also see Pritchard (2016).…”