The euphotic zone constitutes the upper layer of aquatic ecosystems, where ambient light levels are sufficient to support primary production. At any depth within this layer, net primary production (NPP, mg C m −3 d −1 ) rates are always greater than zero (Falkowski, 1994). The base of the euphotic zone (Z eu , m) where the rate of photosynthesis equals that of autotrophic respiration, that is, NPP = 0, has been termed the compensation depth (Z c , m) (Kirk, 1994;Ryther, 1955). Both Z eu and Z c have frequently been used in aquatic ecology as benchmarks for describing and comparing integrated phytoplankton biomass and pigment content, nutrient inventories, NPP, quotients of total water-column light utilization index, and export production processes across different water types and different aquatic ecosystems (Ducklow et al., 2001;