The response of thermospheric composition to geomagnetic storms has been studied for several decades. The first such study was carried out by Seaton (1956), who proposed that an increase in molecular oxygen (O 2 ) number density might account for the decrease of electron density during a major storm that took place on January 25, 194925, . Jacchia (1959 first presented evidence of thermospheric density increases during storms using the measured changes in satellite orbital elements, which were explained in terms of atmospheric heating during geomagnetically disturbed conditions. Prölss and von Zahn (1974aZahn ( , 1974b and Prölss (1980Prölss ( , 1987Prölss ( , 2011 did further studies on the storm-time perturbations in composition, mainly by using the in situ multi-satellite observations near 300 km (e.g., Dynamics Explorer [DE]-2 and ESRO 4 satellites). Based on these studies, a picture of the main thermospheric compositional perturbations during geomagnetic storms was established: an increase of heavier constituents (molecular nitrogen [N 2 ] and oxygen [O 2 ]), and a height-dependent change of atomic oxygen (O) at high-latitudes and middle-latitudes (decreases at lower heights (<∼300 km) and increases higher up); an increase of all constituents at low latitudes over all heights in the thermosphere. In addition, the basic evolution of these perturbations can be described as follows. When a storm begins, strong temperature enhancements occur at high latitudes due to Joule heating, which lead to upward winds (Burns et al., 1995). The nitrogen-rich air is brought up from lower altitudes into the upper thermosphere. The horizontal pressure gradient from temperature differences, in conjunction with ion drag, drives strong horizontal wind changes within a few hours (Ponthieu et al., 1988;Prölss, 2011). Horizontal winds transport the nitrogen-rich air toward the middle-low latitudes at night (