“…This adjustment of roots to increased soil strength reduces the risk of root buckling and decreases the mechanical stress acting on the root during penetration (Materechera et al, 1992;Kirby and Bengough, 2002;Chimungu et al, 2015). Root thickening in response to increased soil strength has been observed in a wide range of species under field and laboratory conditions and often coincides with increased cortical area (Atwell, 1990a;Materechera et al, 1992;Grzesiak et al, 2013;Siczek et al, 2013;Chen et al, 2014;Hernandez-Ramirez et al, 2014;Colombi and Walter, 2016). Since root thickening decreases penetration stress and stabilizes roots, thick roots are likely to be an advantage in soils with increased mechanical impedance (Materechera et al, 1992;Kirby and Bengough, 2002;Chimungu et al, 2015).…”