“…Climate modeling has been able to reconstruct this pattern with very high pCO 2 levels (up to ∼4500 ppm: Huber and Caballero, 2011), but such extremely elevated pCO 2 is not documented by proxy records. It is therefore assumed that Eocene cli-mate sensitivity-often defined as Earth system sensitivity for longer time scales, including both "fast" and "slow" feedbacks (Lunt et al, 2010)-was elevated compared to present, and/ or that other mechanisms, in addition to the dominant forcing of pCO 2 , were in operation (Caballero and Huber, 2013;Anagnostou et al, 2016;Zeebe et al, 2016;Carlson and Caballero, 2016;Cramwinckel et al, 2018;Keery et al, 2018). A variety of geochemical and biological proxies as well as carbon cycle modeling have been used to estimate Eocene pCO 2 , but these estimates still differ hugely (with values ranging from hundreds to thousands of parts per million pCO 2 ); however, there is some convergence forming (Holdgate et al, 2009;Beerling and Royer, 2011;Foster et al, 2017).…”