“…It has been widely applied in systematic and phylogenetic studies providing substantial taxonomic insight (McFadden, France, et al, 2006, McFadden & van Ofwegen 2012, Reijnen et al, 2014, Bryce et al, 2015, Etsabeth, 2018. Some studies have been conducted on S. brassica along the KZN coastline, but these have not fully resolved the taxonomic status of the genus, nor have they looked at population structure Pena-Rodríguez, 2015;Schleyer and Benayahu, 2018;Etsabeth, 2018).…”