“…Many studies had been conducted on biology and sheries of this sh including reproductive biology [9,11,12,13,14,15], population dynamics [11,16], aquaculture [17], taxonomy [4,5,6] and compositions of parasitic organisms [18,19]. However, most of the study on reproductive aspects are concentrated on a particular reproductive trait including length and weight relationship [11,12,13], fecundity [9,14], gonad histology [15], size at rst maturity [11], and size at the rst sex change [12]. Some studies reported also the spawning period of this sh.…”