“…Hovel, 2003;Leibold et al, 2004;Boström et al, 2006). A faunal community living in such patchily distributed habitat has been given to the set of local communities linked by the dispersal of multiple potentially interacting species, that is, the metacommunity (e.g., Hubbell, 2001;Holyoak et al, 2005;Logue et al, 2011;Martiny et al, 2011;Chust et al, 2013a,b. The relative importance of environmental and spatial processes in determining community structures has been evaluated in various local community forms patches in both terrestrial and coastal landscapes, such as ponds, lakes, streams, estuaries, marine pelagic areas, tide pools, and intertidal rocky shores (Cottenie, 2005;Mykrä et al, 2007;Lindo and Winchester, 2009;Pandit et al, 2009;Pinto and MacDougall, 2010;Fiorentino et al, 2012;Chust et al, 2013a). In these studies, the relative contributions of environmental and spatial processes were evaluated in each taxonomic group (e.g., phytoplankton, zooplankton, macro-algae, molluscs), because the relative importance of these two mechanisms varied depending on community characteristics such as ecological traits (e.g., dispersal ability, habitat type, life history, and trophic level).…”