The mesic temperate montane grasslands of the Drakensberg, located within the broader Drakensberg Alpine Centre in southern Africa (Carbutt and Edwards 2003), are productive and ancient (Ellery and Mentis 1992; Bredenkamp et al. 2002; Bond and Parr 2010). Covering more than 42 000 km 2 of the upland area in KwaZulu-Natal (KZN), with only >2 500 km 2 in formal protected areas (Carbutt et al 2011), these closed-canopy grasslands were shaped by conditions during the late Miocene, which facilitated the spread of grassland dominated by C 4 grasses that benefitted from increasing aridity, low atmospheric CO 2 concentrations, and recurrent natural fires (Manry and Knight 1986; Meadows and Linder 1993; Keeley and Rundel 2005; Scheiter et al. 2012). Fire has played an important role in the evolutionary development of the structure and floristic composition of the Drakensberg grassland, with constituent productive grasses that promote fire (Bond et al. 2003) and a diverse assemblage of endemic resprouting forbs with underground bud banks and energy storage units that allow them to withstand frequent burns (Uys 2006; Bond 2016). Fires also maintain the Drakensberg grassland largely free of trees, except for scattered protea species and woody plants in refugia (Adie et al. 2017). A pronounced dry winter together with slow decomposition rates of sourveld grass leaves, result in high loads of cured above-ground biomass (Everson et al. 1988a), which fuel the fires that remove accumulated moribund foliage and litter, allowing sufficient light to reach the soil surface to sustain basal-tillering caespitose grasses (Everson et al. 1985; Everson et al. 1988b; Bond et al. 2003; Neumann et al. 2014) and initiate spring flowering in numerous obligate fire-stimulated grassland forbs (Lamont and Downes 2011). Together, the afore-mentioned appear to indicate a fire regime naturally occurring outside the main growing season of montane grassland plants, but at a time when swards are most susceptible to fire. The frequency and seasonal timing of historical fire regimes in the Drakensberg is unknown and likely variable (Gordijn et al. 2018). Before San hunter-gatherers employed fire to attract wildlife after they settled in the Drakensberg approximately 8 000 to 5 000 BP (Hall 1984; Mazel 1989), lightning was the main natural source of ignition in the